(CNN) — A Manhattan jury has found two Trump Organization companies guilty on multiple charges of criminal tax fraud and falsifying business records connected to a 15-year scheme to defraud tax authorities by failing to report and pay taxes on compensation for top executives.

The Trump Corp. and Trump Payroll Corp. were found guilty on all charges they faced.

Donald Trump and his family were not charged in this case, but the former president was mentioned repeatedly during the trial by prosecutors about his connection to the benefits doled out to certain executives, including company-funded apartments, car leases and personal expenses.

The guilty verdict comes as Trump is under scrutiny by federal and state prosecutors for his handling of classified documents, the effort to overturn the 2020 election results, and the accuracy of the Trump Organization’s business records and financial statements. He is also facing a $250 million civil lawsuit from the New York attorney general alleging he and his adult children were involved in a decade long fraud. The attorney general is seeking to permanently bar them from serving as an officer or director of a company in New York state, among other penalties.

Prosecutor Joshua Steinglass told the jury in closing arguments that Trump “explicitly sanctioned” tax fraud and urged them to reject the defense’s argument that former Trump Organization Chief Financial Officer Allen Weisselberg was a rogue employee motivated by his own personal greed.

“This whole narrative that Donald Trump is blissfully ignorant is just not true,” Steinglass said.

The jury heard that Trump agreed on a whim to pay the private school tuition for his Weisselberg’s grandchildren and signed a lease for a Manhattan apartment to shorten the executive’s commute. Trump personally signed his employees’ bonus checks at Christmas time and he initialed a memo reducing the salary of another top executives, which prosecutors said suggested he knew all along about the fraudulent scheme.

Prosecutors alleged for years top executives reduced their reported salaries by the amount of company-issued fringe benefits to avoid paying the required taxes.

The conviction following the weeks-long trial is a win for the Manhattan district attorney’s office led by Alvin Bragg, which built its case on the Trump Corporation’s general ledger, tax records and the testimony of Weisselberg.

Weisselberg, who is on paid leave from the company, spent three days on the witness stand. He pleaded guilty to 15 felonies for failing to pay taxes on $1.76 million in income. As part of his plea deal, he will be sentenced to five months in jail if the judge finds that he testified truthfully.

In his testimony Weisselberg admitted he should have paid taxes on compensation, totaling roughly $200,000 in one year, which included a luxury Manhattan apartment overlooking the Hudson River, two Mercedes Benz car leases, parking, utilities, furniture and private school tuition for his grandchildren. He also testified that he paid himself and other executives’ bonuses as though they were independent consultants — enabling the Trump companies to evade paying taxes on them.

Weisselberg testified that he pulled off the scheme with the help of his underling, Trump Organization controller Jeffrey McConney. McConney, who received immunity for testifying before the grand jury, admitted to some of the illegal conduct in his testimony.

After Trump was elected president, Weisselberg testified, there was a “clean up” and many of the illegal practices stopped.

He revealed conversations he had with Trump, Eric Trump and Donald Trump Jr., but told the jury when questioned by the Trump attorneys that he did not scheme or conspire with anybody in the Trump family.

Weisselberg became emotional at times, telling the jury he was “embarrassed” by his conduct and that he “betrayed” the Trump family, who has been his employer for 49 years.

Prosecutors and defense attorneys called out Weisselberg’s split loyalties — wanting to live up to his plea deal and serve a lower jail sentence and his loyalty to the Trump family, who could pay him $1 million in compensation this year.

To prove the company’s guilt, prosecutors needed to demonstrate that Weisselberg or McConney was a “high managerial agent” who committed the crimes in the scope of his employment and “in behalf of” the company.

Prosecutors and defense lawyers went in circles arguing over what “in behalf of” meant.

Judge Juan Merchan also struggled over how to explain the phrase to the jury and turned to two legal treaties to fashion a definition.

The judge explained it to the jury, saying, “Under the definition of ‘in behalf of,’ it is not necessary that the criminal acts actually benefit the corporation. But an agent’s acts are not ‘in behalf of’ a corporation if they were undertaken solely to advance the agents own interest. Put another way, if the agent’s acts were taken merely for personal gain, they were not ‘in behalf of’ the corporation.”

Weisselberg walked a fine line in his testimony, telling the jury he never wanted to hurt the company, he was driven by greed and mainly wanted to pay less in taxes. But, he also said, he knew at the time the company would benefit to some degree from his schemes.

In his testimony Weisselberg said, “It was a benefit to the company but primarily it was due to my greed.” He told the jury that the company saved money by paying less taxes on his off-the-books compensation and acknowledged when asked by prosecutor Susan Hoffinger if, while his primary goal was to avoid taxes, it also created a benefit to the company.

“To some degree, yes,” Weisselberg testified.

Weisselberg said he and McConney knew at the time that the company would pay less payroll taxes through the scheme although he said they never discussed it explicitly.

The Trump attorneys argued, repeatedly, to the jury that “Weisselberg did it for Weisselberg” to emphasize that he was motivated solely by his personal greed.

On cross examination, Weisselberg agreed that the decision to not pay taxes was his and made solely to benefit himself.

“That was my intent,” Weisselberg said when questioned by the Trump attorneys, “to benefit myself.”

The lawyers for the Trump entities called just one fact witness, the real estate companies’ long-time accountant Donald Bender from Mazars USA, which dropped Trump as a client earlier this year.

Trump attorneys said Bender was either aware of the off-the-books compensation or should have caught the tax fraud and they accused him of lying on the stand.

When questioned by prosecutors Bender testified that he trusted and relied on Weisselberg, who testified that he hid the illegal scheme.

Steinglass, the prosecutor, told the jury the Trump companies were guilty, and the illegal scheme was concocted “so the employees can get more net pay while costing the Trump Corporation less. It’s a win-win — unless you’re the tax authorities.”

This story has been updated with additional details.